To correct for the number of traits being tested per tissue region, a false discovery rate (FDR) threshold was determined based on the empirical p-values using the fwer2fdr function of the multtest package in R version 2.6.1. Empirical p-values were allowed to exceed this threshold if their linkage disequilibrium r2 was greater than or equal to 0.7 with a SNP with empirical values within the FDR threshold.
