We divided the trans-QTLs into two broad subgroups—those with peak QTLs on markers in the proximal part of Qrr1 (Qrr1p; 172–174.5 Mb or segments 1, 2, 3 in figure 2), and those with peak QTLs on markers in the distal part of Qrr1 (Qrr1d; 174.5–177.5 Mb or segments 4, 5, and 6 in figure 2). While Qrr1p is relatively gene-rich, only 35% of the trans-QTLs (129 out of 365 probe sets) have peak LOD scores in this region. The majority of trans-QTLs—about 65% (236 out of 365 probe sets)—have peak QTLs in the relatively gene-sparse Qrr1d.