conditional analysis 59 was used to identify independently-segregating variants in each of the merged loci that was genome-wide significant, using the linkage disequilibrium structure of UK Biobank as a reference panel. Genome-wide statistical significance was defined as P < 5 × 10-8, with the meta-analysis results from Metal reported. Due to the complexity of the major histocompatibility complex region only the most significant variant in that region is reported.