and with hypotheses that global, genetically mediated differences in cell division were the driving force behind interspecies differences in total brain volume (Darlington et al., 1999; Finlay and Darlington, 1995; Fishell, 1997). Expanding the entire brain when only specific functions might be selected for is metabolically costly, but the number of mutations required to affect cell division would be far less than that required to completely change cerebral organization.