In addition, we found a significant fraction of cells with increased but not decreased theta-band phase coupling during contraversive as compared to ipsiversive trials (ACC units: 34/353=9.6%, P<0.001; FEF units: 15/209=7.2%, P<0.001, Binomial test; Fig. 5a,b, right). Furthermore, we observed that the average spike-LFP phase locking was increased during contraversive as compared to ipsiversive trials in a subset of theta frequencies (3–4 Hz; Fig. 6a,b, right).