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Chunk #44 — Model — Genotypic Values — Complementary Model

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Data and theory point to mainly additive genetic variance for complex traits.
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Another simple epistatic model involving all components is the following:which can also be defined for multiple loci. For two loci, for example, using similar methods it can be shown that: for pi = 0.5, V A/V G = 0.56; with the uniform distribution, E(V A)/E(V G) = 2/3; and with the ‘U’ distribution .