involved in behavioral control: a model-based (goal-directed) system that may involve a number of cortical areas in addition to parts of anterior medial striatum, and a model-free (habitual) system that may be depend predominantly on dopaminergic-striatal pathways (Balleine et al., 2007). However, the data presented here, in particular the weighted combination of both models in the hybrid account, suggests that both learning mechanisms act together to produce effective action selection rather than dualistic processing of two learning modules that exert separate control over choice behavior.