NLS designed and carried out the statistical analyses and power calculations and drafted the manuscript. SFS carried out SNP selection for genotyping and contributed to the design of the study. AMG contributed to the conception and design of the study. RAG carried out the population structure analyses. ALH performed linkage disequilibrium analyses and visualization. JPR contributed to the conception and design of the study and advised on the implementation of the method. LJB contributed to the conception and design of the study and is the PI of the FSCD project that provided the samples. All authors contributed to the interpretation of results and the intellectual content of the manuscript, and have read and approved the final manuscript.