less than 0.1 ms (Schmitzer-Torbert and Redish, 2008), as well as a waveform peak width smaller than 0.15 ms, waveform valley width smaller than 0.35 ms, and a firing rate above 2 Hz (adapted from Berke et al., 2004) were considered (Figure 1). Additionally, only cells with L-ratio (a measure of isolation quality, Schmitzer-Torbert et al., 2005) ≤0.1 and average peak amplitude ≥80 μV were included for analysis. We did not make attempts to distinguish neurons that were recorded for multiple recording sessions, so inferences about the frequency or distribution of certain properties over this population should be made with this potentially uneven sampling in mind.