The hippocampal coupling account predicts that HF regions will show increased activity, together with the DMN regions during successful retrieval regardless of internal/external orientation. As noted, in line with this prediction (see Table 1), all DMN regions including HF showed a significant main effect of memory, and no memory×orientation interaction (Table 4, Figure 2). The hippocampal coupling account also predicts that HF regions will show increased activity during successful encoding, but the DMN regions decreased activity, regardless of internal/external orientation. In line with this prediction, as illustrated in Table 3 and Figure 2, we found that the DMN regions, except for superior PFC, showed a negative main effect of memory, whereas HF showed a positive main effect of memory. None of these regions showed a significant interaction effect between memory and orientation during encoding. Overall, these findings provide clear support for the hippocampal coupling, but not for the internal orienting, hypothesis.
