Data were analyzed using SPM5 (Wellcome Department of Imaging Neuroscience, University College, London). Functional volumes were corrected for differences in slice acquisition timing and rigid-body realigned to the initial volume of the first functional imaging scan to account for residual movement after prospective motion correction (Thesen et al., 2000). Each subject's high resolution anatomic image was co-registered to the reference functional volume, segmented into gray, white and cerebrospinal fluid tissue components. Nonlinear spatial transformation parameters from this segmentation were subsequently applied to transform functional image volumes into the Montreal Neurological Institute (MNI) coordinate space, which were re-sampled to isotropic 2 mm voxels and smoothed by a 6 mm full-width at half-maximum (FWHM) isotropic Gaussian kernel.