the trait of interest (as opposed to fitness itself) is not a requirement of these latter forms of balancing selection (Turelli and Barton 2004), and since additive genetic variation is maintained (rather than depleted) by balancing selection, a high proportion of nonadditive variation is not expected. Furthermore, there would be no reason to expect inbreeding to affect the trait since that requires directional dominance with respect to the trait (Charlesworth and Charlesworth 1987; Roff 2005).
