e=(pA22pA(1−pA)(1−pA)2pApEpA(1−pE)+(1−pA)pE(1−pA)(1−pE)pE22pE(1−pE)(1−pE)2) where for clarity we have dropped the variable t indicating the locus, and we set the transmission probabilities aij for i,j=0..2, where i,j indicate the number of chromosomes of European ancestry at a given locus and the next locus, as
