R.G., C.F., J.-J.R., T.R.K., and J.C. conceived and designed research; R.G., C.M., and M.W. performed experiments; R.G., T.R.K., and J.C. analyzed data; R.G., T.R.K., and J.C. interpreted results of experiments; R.G. and J.C. prepared figures; R.G. drafted manuscript; R.G., C.M., M.W., C.F., J.-J.R., T.R.K., and J.C. edited and revised manuscript; R.G., C.M., M.W., C.F., J.-J.R., T.R.K., and J.C. approved final version of manuscript.