SNP-based P-values from the meta-analysis were used as input for the gene-based genome-wide association analysis (GWGAS). 18,128 protein-coding genes (each containing at least 1 GWAS SNP) from the NCBI 37.3 gene definitions were used as basis for GWGAS in MAGMA25 (see URLs). The Bonferroni-corrected genome-wide significance threshold was .05/18,128 genes=2.76×10-6.