no overt morphological or behavioral phenotypes. Heads of homozygous dallyMB950 flies were shown to have a 27% reduction in dally transcript (P = 0.0218, n = 5 replicates, two-sample t-test), but dallyMB950 complemented the sterility and wing vein morphology phenotypes of the strong loss-of-function allele dally80. Given the molecular evidence that dallyMB950 is an allele of dally, a transposon inserted into the gene coupled with reduced transcript production, we conclude that intragenic complementation is occurring because the dallyMB950 allele provides enough function to complement the phenotype of the strong loss-of-function allele.