coincidence of the WHR signal with expression for rs984222 (TBX15 in omental fat), rs1055144 (expressed sequence tag AA553656 in SAT), rs10195252 (GRB14 in SAT), rs4823006 (ZNRF3 in SAT and omental fat), and rs6784615 (STAB1 in blood)(Table 4). Taken together, the overlap between trait association and gene expression at these loci suggests that the WHR associations are driven through altered expression of PIGC, TBX15, AA553656, GRB1, ZNRF3 and STAB1.