Drosophila strains were maintained on standard cornmeal/molasses/yeast media at 25°C and 70% humidity with an approximately 16-hr/8-hr light/dark schedule. All strains were outcrossed for five generations to the Berlin genetic background strain carrying the w1118 eye-color marker. Strain sources were dallyMB950 (MB00950: Bloomington Stock Center), dally80 (Xinhua Lin), dlpf03537 (Exelixis/Harvard), and dlp1 (Scott Selleck).