Second, we plan to test the hypothesis that individual differences in the function, structure, and connectivity of these same four neural hubs are correlated with real-world behaviours needed to prepare for successful ageing. We will focus on neural correlates of midlife real-world behaviours outside the lab (i.e. “neuroecology”) that prepare individuals for health and wellbeing in late life, such as preparing for the financial future, building social relationships, and sustaining positive emotions.