prefrontal cortex, visual cortex and cerebellum, respectively66, liver58,67–70, osteoblasts71, intestine72, lung73, skin50,74 and primary fibroblasts48. Micro-RNA QTLs were also queried for gluteal and abdominal adipose75. Only results that reach study-wise significance thresholds in their respective datasets were included (Supplementary table 6). Expression data was also available on adipose tissue and whole blood samples from deCODE where parent-of-origin specific analyses were possible.