Path coefficients of the multivariate Cholesky decomposition can be found in Figure 1 and results are summarised in Table 5. To test the significance of each path in the Cholesky decomposition, genetic and environmental parameters were dropped from the model and model fit was compared with the full model using an α-level of 0.05. Because the parameter estimates here are based on multivariate relationships, the estimates may differ from the estimates obtained in the univariate models and the expectations from the bivariate phenotypic correlations presented in Table 4.